Reversion of fruit-dependent inhibition of flowering in Citrusrequires sprouting of buds with epigenetically silenced CcMADS19

In Citrus, the response to environmental floral inductive signals is inhibited by the presence of developing fruits. The mechanism involves epigenetic activation of the CcMADS19 locus (FLC orthologue), encoding a floral repressor. To understand how this epigenetic regulation is reverted to allow flo...

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Autores principales: Mesejo, Carlos, Marzal, Andrés, Martínez-Fuentes, Amparo, Reig, Carmina, De-Lucas, Miguel, Iglesias, Domingo J., Primo-Millo, Eduardo, Blazquez, Miguel A., Agustí, Manuel
Formato: article
Lenguaje:Inglés
Publicado: Wiley Online Library 2021
Materias:
Acceso en línea:http://hdl.handle.net/20.500.11939/7659
https://nph.onlinelibrary.wiley.com/doi/full/10.1111/nph.17681
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author Mesejo, Carlos
Marzal, Andrés
Martínez-Fuentes, Amparo
Reig, Carmina
De-Lucas, Miguel
Iglesias, Domingo J.
Primo-Millo, Eduardo
Blazquez, Miguel A.
Agustí, Manuel
author_browse Agustí, Manuel
Blazquez, Miguel A.
De-Lucas, Miguel
Iglesias, Domingo J.
Martínez-Fuentes, Amparo
Marzal, Andrés
Mesejo, Carlos
Primo-Millo, Eduardo
Reig, Carmina
author_facet Mesejo, Carlos
Marzal, Andrés
Martínez-Fuentes, Amparo
Reig, Carmina
De-Lucas, Miguel
Iglesias, Domingo J.
Primo-Millo, Eduardo
Blazquez, Miguel A.
Agustí, Manuel
author_sort Mesejo, Carlos
collection ReDivia
description In Citrus, the response to environmental floral inductive signals is inhibited by the presence of developing fruits. The mechanism involves epigenetic activation of the CcMADS19 locus (FLC orthologue), encoding a floral repressor. To understand how this epigenetic regulation is reverted to allow flowering in the following season, we have forced precocious sprouting of axillary buds in fruit-bearing shoots, and examined the competence to floral inductive signals of old and new leaves derived from them. We have found that CcMADS19 is enriched in repressive H3K27me3 marks in young, but not old leaves, revealing that axillary buds retain a silenced version of the floral repressor that is mitotically transmitted to the newly emerging leaves, which are able to induce flowering. Therefore, we propose that flowering in Citrus is necessarily preceded by vegetative sprouting, so that the competence to respond to floral inductive signals is reset in the new leaves.
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spelling ReDivia76592025-04-25T14:48:27Z Reversion of fruit-dependent inhibition of flowering in Citrusrequires sprouting of buds with epigenetically silenced CcMADS19 Mesejo, Carlos Marzal, Andrés Martínez-Fuentes, Amparo Reig, Carmina De-Lucas, Miguel Iglesias, Domingo J. Primo-Millo, Eduardo Blazquez, Miguel A. Agustí, Manuel Alternate bearing Chromatin remodelling FLC (Flowering locus C) FT (Flowering locus T) F62 Plant physiology - Growth and development F63 Plant physiology - Reproduction Citrus Flowering In Citrus, the response to environmental floral inductive signals is inhibited by the presence of developing fruits. The mechanism involves epigenetic activation of the CcMADS19 locus (FLC orthologue), encoding a floral repressor. To understand how this epigenetic regulation is reverted to allow flowering in the following season, we have forced precocious sprouting of axillary buds in fruit-bearing shoots, and examined the competence to floral inductive signals of old and new leaves derived from them. We have found that CcMADS19 is enriched in repressive H3K27me3 marks in young, but not old leaves, revealing that axillary buds retain a silenced version of the floral repressor that is mitotically transmitted to the newly emerging leaves, which are able to induce flowering. Therefore, we propose that flowering in Citrus is necessarily preceded by vegetative sprouting, so that the competence to respond to floral inductive signals is reset in the new leaves. 2021-10-13T10:51:05Z 2021-10-13T10:51:05Z 2021 article publishedVersion Mesejo, C., Marzal, A., Martínez‐Fuentes, A., Reig, C., de-Lucas, M., Iglesias, D. J. et al. (2021). Reversion of fruit‐dependent inhibition of flowering in Citrus requires sprouting of buds with epigenetically silenced CcMADS19. New Phytologist, 2021, 17681. 1469-8137 0028-646X (e-ISSN) http://hdl.handle.net/20.500.11939/7659 10.1111/nph.17681 https://nph.onlinelibrary.wiley.com/doi/full/10.1111/nph.17681 en info:eu-repo/grantAgreement/GVA/PROMETEO 2019/021 info:eu-repo/grantAgreement/MICINN//RTI2018-101303-B-100 The work described in this paper was partially supported by grants PROMETEO/2019/021 Generalitat valenciana (MAB). Financial supported also by the Spanish Ministerio de Ciencia, Innovación y Universidades through grant nº RTI2018-101303-B-100 (CM). Atribución-NoComercial-SinDerivadas 3.0 España http://creativecommons.org/licenses/by-nc-nd/3.0/es/ openAccess Wiley Online Library electronico
spellingShingle Alternate bearing
Chromatin remodelling
FLC (Flowering locus C)
FT (Flowering locus T)
F62 Plant physiology - Growth and development
F63 Plant physiology - Reproduction
Citrus
Flowering
Mesejo, Carlos
Marzal, Andrés
Martínez-Fuentes, Amparo
Reig, Carmina
De-Lucas, Miguel
Iglesias, Domingo J.
Primo-Millo, Eduardo
Blazquez, Miguel A.
Agustí, Manuel
Reversion of fruit-dependent inhibition of flowering in Citrusrequires sprouting of buds with epigenetically silenced CcMADS19
title Reversion of fruit-dependent inhibition of flowering in Citrusrequires sprouting of buds with epigenetically silenced CcMADS19
title_full Reversion of fruit-dependent inhibition of flowering in Citrusrequires sprouting of buds with epigenetically silenced CcMADS19
title_fullStr Reversion of fruit-dependent inhibition of flowering in Citrusrequires sprouting of buds with epigenetically silenced CcMADS19
title_full_unstemmed Reversion of fruit-dependent inhibition of flowering in Citrusrequires sprouting of buds with epigenetically silenced CcMADS19
title_short Reversion of fruit-dependent inhibition of flowering in Citrusrequires sprouting of buds with epigenetically silenced CcMADS19
title_sort reversion of fruit dependent inhibition of flowering in citrusrequires sprouting of buds with epigenetically silenced ccmads19
topic Alternate bearing
Chromatin remodelling
FLC (Flowering locus C)
FT (Flowering locus T)
F62 Plant physiology - Growth and development
F63 Plant physiology - Reproduction
Citrus
Flowering
url http://hdl.handle.net/20.500.11939/7659
https://nph.onlinelibrary.wiley.com/doi/full/10.1111/nph.17681
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