Absence of the nitrous oxide reductase gene cluster in commercial alfalfa inoculants is probably due to the extensive loss of genes during rhizobial domestication

As other legume crops, alfalfa cultivation increases the emission of the greenhouse gas nitrous oxide (N2O). Since legumesymbiotic nitrogen-fixing bacteria play a crucial role in this emission, it is important to understand the possible impacts of rhizobial domestication on the evolution of denitr...

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Main Authors: Brambilla, Silvina Maricel, Frare, Romina Alejandra, Soto, Gabriela Cinthia, Jozefkowicz, Cintia, Ayub, Nicolás Daniel
Format: info:ar-repo/semantics/artículo
Language:Inglés
Published: Springer 2018
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Online Access:https://link.springer.com/article/10.1007/s00248-018-1145-9
http://hdl.handle.net/20.500.12123/3399
https://doi.org/10.1007/s00248-018-1145-9
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author Brambilla, Silvina Maricel
Frare, Romina Alejandra
Soto, Gabriela Cinthia
Jozefkowicz, Cintia
Ayub, Nicolás Daniel
author_browse Ayub, Nicolás Daniel
Brambilla, Silvina Maricel
Frare, Romina Alejandra
Jozefkowicz, Cintia
Soto, Gabriela Cinthia
author_facet Brambilla, Silvina Maricel
Frare, Romina Alejandra
Soto, Gabriela Cinthia
Jozefkowicz, Cintia
Ayub, Nicolás Daniel
author_sort Brambilla, Silvina Maricel
collection INTA Digital
description As other legume crops, alfalfa cultivation increases the emission of the greenhouse gas nitrous oxide (N2O). Since legumesymbiotic nitrogen-fixing bacteria play a crucial role in this emission, it is important to understand the possible impacts of rhizobial domestication on the evolution of denitrification genes. In comparison with the genomes of non-commercial strains, those of commercial alfalfa inoculants exhibit low total genome size, low number of ORFs and high numbers of both frameshifted genes and pseudogenes, suggesting a dramatic loss of genes during bacterial domestication. Genomic analysis focused on denitrification genes revealed that commercial strains have perfectly conserved the nitrate (NAP), nitrite (NIR) and nitric (NOR) reductase clusters related to the production of N2O from nitrate but completely lost the nitrous oxide (NOS) reductase cluster (nosRZDFYLX genes) associated with the reduction of N2O to gas nitrogen. Based on these results, we propose future screenings for alfalfa-nodulating isolates containing both nitrogen fixation and N2O reductase genes for environmental sustainability of alfalfa production.
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institution Instituto Nacional de Tecnología Agropecuaria (INTA -Argentina)
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spelling INTA33992018-09-19T17:10:15Z Absence of the nitrous oxide reductase gene cluster in commercial alfalfa inoculants is probably due to the extensive loss of genes during rhizobial domestication Brambilla, Silvina Maricel Frare, Romina Alejandra Soto, Gabriela Cinthia Jozefkowicz, Cintia Ayub, Nicolás Daniel Óxido Nitroso Medicago sativa Fijación del Nitrógeno Muestreo cluster Genes Rhizobium Nitrous Oxide Nitrogen Fixation Cluster Sampling Inoculantes comerciales As other legume crops, alfalfa cultivation increases the emission of the greenhouse gas nitrous oxide (N2O). Since legumesymbiotic nitrogen-fixing bacteria play a crucial role in this emission, it is important to understand the possible impacts of rhizobial domestication on the evolution of denitrification genes. In comparison with the genomes of non-commercial strains, those of commercial alfalfa inoculants exhibit low total genome size, low number of ORFs and high numbers of both frameshifted genes and pseudogenes, suggesting a dramatic loss of genes during bacterial domestication. Genomic analysis focused on denitrification genes revealed that commercial strains have perfectly conserved the nitrate (NAP), nitrite (NIR) and nitric (NOR) reductase clusters related to the production of N2O from nitrate but completely lost the nitrous oxide (NOS) reductase cluster (nosRZDFYLX genes) associated with the reduction of N2O to gas nitrogen. Based on these results, we propose future screenings for alfalfa-nodulating isolates containing both nitrogen fixation and N2O reductase genes for environmental sustainability of alfalfa production. Instituto de Genética Fil: Brambilla, Silvina Maricel. Instituto Nacional de Tecnología Agropecuaria (INTA). Instituto de Genética "Ewald A. Favret"; Argentina Fil: Frare, Romina Alejandra. Instituto Nacional de Tecnología Agropecuaria (INTA). Instituto de Genética "Ewald A. Favret"; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina Fil: Soto, Gabriela Cinthia. Instituto Nacional de Tecnología Agropecuaria (INTA). Instituto de Genética "Ewald A. Favret"; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina Fil: Jozefkowicz, Cintia. Instituto Nacional de Tecnología Agropecuaria (INTA). Instituto de Genética "Ewald A. Favret"; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina Fil: Ayub, Nicolás Daniel. Instituto Nacional de Tecnología Agropecuaria (INTA). Instituto de Genética "Ewald A. Favret"; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina 2018-09-19T17:04:53Z 2018-09-19T17:04:53Z 2018-08 info:ar-repo/semantics/artículo info:eu-repo/semantics/article info:eu-repo/semantics/publishedVersion https://link.springer.com/article/10.1007/s00248-018-1145-9 http://hdl.handle.net/20.500.12123/3399 0095-3628 1432-184X https://doi.org/10.1007/s00248-018-1145-9 eng info:eu-repo/semantics/restrictedAccess application/pdf Springer Microbial Ecology 76 (2): 299–302 (August 2018)
spellingShingle Óxido Nitroso
Medicago sativa
Fijación del Nitrógeno
Muestreo cluster
Genes
Rhizobium
Nitrous Oxide
Nitrogen Fixation
Cluster Sampling
Inoculantes comerciales
Brambilla, Silvina Maricel
Frare, Romina Alejandra
Soto, Gabriela Cinthia
Jozefkowicz, Cintia
Ayub, Nicolás Daniel
Absence of the nitrous oxide reductase gene cluster in commercial alfalfa inoculants is probably due to the extensive loss of genes during rhizobial domestication
title Absence of the nitrous oxide reductase gene cluster in commercial alfalfa inoculants is probably due to the extensive loss of genes during rhizobial domestication
title_full Absence of the nitrous oxide reductase gene cluster in commercial alfalfa inoculants is probably due to the extensive loss of genes during rhizobial domestication
title_fullStr Absence of the nitrous oxide reductase gene cluster in commercial alfalfa inoculants is probably due to the extensive loss of genes during rhizobial domestication
title_full_unstemmed Absence of the nitrous oxide reductase gene cluster in commercial alfalfa inoculants is probably due to the extensive loss of genes during rhizobial domestication
title_short Absence of the nitrous oxide reductase gene cluster in commercial alfalfa inoculants is probably due to the extensive loss of genes during rhizobial domestication
title_sort absence of the nitrous oxide reductase gene cluster in commercial alfalfa inoculants is probably due to the extensive loss of genes during rhizobial domestication
topic Óxido Nitroso
Medicago sativa
Fijación del Nitrógeno
Muestreo cluster
Genes
Rhizobium
Nitrous Oxide
Nitrogen Fixation
Cluster Sampling
Inoculantes comerciales
url https://link.springer.com/article/10.1007/s00248-018-1145-9
http://hdl.handle.net/20.500.12123/3399
https://doi.org/10.1007/s00248-018-1145-9
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