An empirical model that uses light attenuation and plant nitrogen status to predict within-canopy N distribution and upscale photosynthesis from leaf to whole canopy

Modelling the spatial and temporal distribution of leaf nitrogen (N) is central to specify photosynthetic parameters and simulate canopy photosynthesis. Leaf photosynthetic parameters depend on both local light availability and whole-plant N status. The interaction between these two levels of inte...

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Autores principales: Louarn, Gaëtan, Frak, Ela, Zaka, Serge, Prieto, Jorge Alejandro, Lebon, Eric
Formato: Artículo
Lenguaje:Inglés
Publicado: 2017
Materias:
Acceso en línea:http://hdl.handle.net/20.500.12123/1319
https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4635319/
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author Louarn, Gaëtan
Frak, Ela
Zaka, Serge
Prieto, Jorge Alejandro
Lebon, Eric
author_browse Frak, Ela
Lebon, Eric
Louarn, Gaëtan
Prieto, Jorge Alejandro
Zaka, Serge
author_facet Louarn, Gaëtan
Frak, Ela
Zaka, Serge
Prieto, Jorge Alejandro
Lebon, Eric
author_sort Louarn, Gaëtan
collection INTA Digital
description Modelling the spatial and temporal distribution of leaf nitrogen (N) is central to specify photosynthetic parameters and simulate canopy photosynthesis. Leaf photosynthetic parameters depend on both local light availability and whole-plant N status. The interaction between these two levels of integration has generally been modelled by assuming optimal canopy functioning, which is not supported by experiments. During this study, we examined how a set of empirical relationships with measurable parameters could be used instead to predict photosynthesis at the leaf and whole-canopy levels. The distribution of leaf N per unit area (Na) within the canopy was related to leaf light irradiance and to the nitrogen nutrition index (NNI), a whole-plant variable accounting for plant N status. Na was then used to determine the photosynthetic parameters of a leaf gas exchange model. The model was assessed on alfalfa canopies under contrasting N nutrition and with N2-fixing and non-fixing plants. Three experiments were carried out to parameterize the relationships between Na, leaf irradiance, NNI and photosynthetic parameters. An additional independent data set was used for model evaluation. The N distribution model showed that it was able to predict leaf N on the set of leaves tested. The Na at the top of the canopy appeared to be related linearly to the NNI, whereas the coef- ficient accounting for N allocation remained constant. Photosynthetic parameters were related linearly to Na irrespective of N nutrition and the N acquisition mode. Daily patterns of gas exchange were simulated accurately at the leaf scale. When integrated at the whole-canopy scale, the model predicted that raising N availability above an NNI of 1 did not result in increased net photosynthesis. Overall, the model proposed offered a solution for a dynamic coupling of leaf photosynthesis and canopy N distribution without requiring any optimal functioning hypothesis.
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spelling INTA13192018-01-12T13:49:57Z An empirical model that uses light attenuation and plant nitrogen status to predict within-canopy N distribution and upscale photosynthesis from leaf to whole canopy Louarn, Gaëtan Frak, Ela Zaka, Serge Prieto, Jorge Alejandro Lebon, Eric Medicago Sativa Cubierta Vegetal Fotosíntesis Nitrógeno Disponibilidad de Nutrientes Transpiración Superficie Foliar Modelos Cubierta de Copas Plant Cover Photosynthesis Nitrogen Nutrient Availability Transpiration Leaf Area Models Canopy Nitrogen Nutrition Index Alfalfa Canipia Modelling the spatial and temporal distribution of leaf nitrogen (N) is central to specify photosynthetic parameters and simulate canopy photosynthesis. Leaf photosynthetic parameters depend on both local light availability and whole-plant N status. The interaction between these two levels of integration has generally been modelled by assuming optimal canopy functioning, which is not supported by experiments. During this study, we examined how a set of empirical relationships with measurable parameters could be used instead to predict photosynthesis at the leaf and whole-canopy levels. The distribution of leaf N per unit area (Na) within the canopy was related to leaf light irradiance and to the nitrogen nutrition index (NNI), a whole-plant variable accounting for plant N status. Na was then used to determine the photosynthetic parameters of a leaf gas exchange model. The model was assessed on alfalfa canopies under contrasting N nutrition and with N2-fixing and non-fixing plants. Three experiments were carried out to parameterize the relationships between Na, leaf irradiance, NNI and photosynthetic parameters. An additional independent data set was used for model evaluation. The N distribution model showed that it was able to predict leaf N on the set of leaves tested. The Na at the top of the canopy appeared to be related linearly to the NNI, whereas the coef- ficient accounting for N allocation remained constant. Photosynthetic parameters were related linearly to Na irrespective of N nutrition and the N acquisition mode. Daily patterns of gas exchange were simulated accurately at the leaf scale. When integrated at the whole-canopy scale, the model predicted that raising N availability above an NNI of 1 did not result in increased net photosynthesis. Overall, the model proposed offered a solution for a dynamic coupling of leaf photosynthesis and canopy N distribution without requiring any optimal functioning hypothesis. Fil: Louarn, Gaëtan. Institut National de la Recherche Agronomique; Francia Fil: Frak, Ela. Institut National de la Recherche Agronomique; Francia Fil: Zaka, Serge. Institut National de la Recherche Agronomique; Francia Fil: Lebon, Eric. Institut National de la Recherche Agronomique. Unité Mixte de Recherche; Francia Fil: Prieto, Jorge Alejandro. Instituto Nacional de Tecnología Agropecuaria (INTA). Estación Experimental Agropecuaria Mendoza; Argentina 2017-09-25T17:55:34Z 2017-09-25T17:55:34Z 2015-10-03 info:ar-repo/semantics/artículo info:eu-repo/semantics/article info:eu-repo/semantics/publishedVersion http://hdl.handle.net/20.500.12123/1319 https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4635319/ 2041-2851 eng info:eu-repo/semantics/openAccess http://creativecommons.org/licenses/by-nc-sa/4.0/ Creative Commons Attribution-NonCommercial-ShareAlike 4.0 International (CC BY-NC-SA 4.0) application/pdf AoB plants 7 : 1-16. (2015)
spellingShingle Medicago Sativa
Cubierta Vegetal
Fotosíntesis
Nitrógeno
Disponibilidad de Nutrientes
Transpiración
Superficie Foliar
Modelos
Cubierta de Copas
Plant Cover
Photosynthesis
Nitrogen
Nutrient Availability
Transpiration
Leaf Area
Models
Canopy
Nitrogen Nutrition Index
Alfalfa
Canipia
Louarn, Gaëtan
Frak, Ela
Zaka, Serge
Prieto, Jorge Alejandro
Lebon, Eric
An empirical model that uses light attenuation and plant nitrogen status to predict within-canopy N distribution and upscale photosynthesis from leaf to whole canopy
title An empirical model that uses light attenuation and plant nitrogen status to predict within-canopy N distribution and upscale photosynthesis from leaf to whole canopy
title_full An empirical model that uses light attenuation and plant nitrogen status to predict within-canopy N distribution and upscale photosynthesis from leaf to whole canopy
title_fullStr An empirical model that uses light attenuation and plant nitrogen status to predict within-canopy N distribution and upscale photosynthesis from leaf to whole canopy
title_full_unstemmed An empirical model that uses light attenuation and plant nitrogen status to predict within-canopy N distribution and upscale photosynthesis from leaf to whole canopy
title_short An empirical model that uses light attenuation and plant nitrogen status to predict within-canopy N distribution and upscale photosynthesis from leaf to whole canopy
title_sort empirical model that uses light attenuation and plant nitrogen status to predict within canopy n distribution and upscale photosynthesis from leaf to whole canopy
topic Medicago Sativa
Cubierta Vegetal
Fotosíntesis
Nitrógeno
Disponibilidad de Nutrientes
Transpiración
Superficie Foliar
Modelos
Cubierta de Copas
Plant Cover
Photosynthesis
Nitrogen
Nutrient Availability
Transpiration
Leaf Area
Models
Canopy
Nitrogen Nutrition Index
Alfalfa
Canipia
url http://hdl.handle.net/20.500.12123/1319
https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4635319/
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