A nomadic subtelomeric disease resistance gene cluster in common bean

The B4 resistance (R) gene cluster is one of the largest clusters known in common bean (Phaseolus vulgaris [Pv]). It is located in a peculiar genomic environment in the subtelomeric region of the short arm of chromosome 4, adjacent to two heterochromatic blocks (knobs). We sequenced 650 kb spanning...

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Autores principales: David, Perrine, Chen, Nicolas W.G., Pedrosa Harand, Andrea, Thareau, Vincent, Sévignac, Mireille, Cannon, Steven B., Debouck, Daniel G., Langin, Thierry, Geffroy, Valerie
Formato: Journal Article
Lenguaje:Inglés
Publicado: Oxford University Press 2009
Materias:
Acceso en línea:https://hdl.handle.net/10568/83269
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author David, Perrine
Chen, Nicolas W.G.
Pedrosa Harand, Andrea
Thareau, Vincent
Sévignac, Mireille
Cannon, Steven B.
Debouck, Daniel G.
Langin, Thierry
Geffroy, Valerie
author_browse Cannon, Steven B.
Chen, Nicolas W.G.
David, Perrine
Debouck, Daniel G.
Geffroy, Valerie
Langin, Thierry
Pedrosa Harand, Andrea
Sévignac, Mireille
Thareau, Vincent
author_facet David, Perrine
Chen, Nicolas W.G.
Pedrosa Harand, Andrea
Thareau, Vincent
Sévignac, Mireille
Cannon, Steven B.
Debouck, Daniel G.
Langin, Thierry
Geffroy, Valerie
author_sort David, Perrine
collection Repository of Agricultural Research Outputs (CGSpace)
description The B4 resistance (R) gene cluster is one of the largest clusters known in common bean (Phaseolus vulgaris [Pv]). It is located in a peculiar genomic environment in the subtelomeric region of the short arm of chromosome 4, adjacent to two heterochromatic blocks (knobs). We sequenced 650 kb spanning this locus and annotated 97 genes, 26 of which correspond to Coiled-Coil-Nucleotide-Binding-Site-Leucine-Rich-Repeat (CNL). Conserved microsynteny was observed between the Pv B4 locus and corresponding regions of Medicago truncatula and Lotus japonicus in chromosomes Mt6 and Lj2, respectively. The notable exception was the CNL sequences, which were completely absent in these regions. The origin of the Pv B4-CNL sequences was investigated through phylogenetic analysis, which reveals that, in the Pv genome, paralogous CNL genes are shared among nonhomologous chromosomes (4 and 11). Together, our results suggest that Pv B4-CNL was derived from CNL sequences from another cluster, the Co-2 cluster, through an ectopic recombination event. Integration of the soybean (Glycine max) genome data enables us to date more precisely this event and also to infer that a single CNL moved from the Co-2 to the B4 cluster. Moreover, we identified a new 528-bp satellite repeat, referred to as khipu, specific to the Phaseolus genus, present both between B4-CNL sequences and in the two knobs identified at the B4 R gene cluster. The khipu repeat is present on most chromosomal termini, indicating the existence of frequent ectopic recombination events in Pv subtelomeric regions. Our results highlight the importance of ectopic recombination in R gene evolution.
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spelling CGSpace832692025-11-12T05:57:23Z A nomadic subtelomeric disease resistance gene cluster in common bean David, Perrine Chen, Nicolas W.G. Pedrosa Harand, Andrea Thareau, Vincent Sévignac, Mireille Cannon, Steven B. Debouck, Daniel G. Langin, Thierry Geffroy, Valerie phaseolus vulgaris phylogeny recombination nucleotide sequence chromosomes disease resistance filogenia recombinación secuencia nucleotidica cromosomas resistencia a la enfermedad The B4 resistance (R) gene cluster is one of the largest clusters known in common bean (Phaseolus vulgaris [Pv]). It is located in a peculiar genomic environment in the subtelomeric region of the short arm of chromosome 4, adjacent to two heterochromatic blocks (knobs). We sequenced 650 kb spanning this locus and annotated 97 genes, 26 of which correspond to Coiled-Coil-Nucleotide-Binding-Site-Leucine-Rich-Repeat (CNL). Conserved microsynteny was observed between the Pv B4 locus and corresponding regions of Medicago truncatula and Lotus japonicus in chromosomes Mt6 and Lj2, respectively. The notable exception was the CNL sequences, which were completely absent in these regions. The origin of the Pv B4-CNL sequences was investigated through phylogenetic analysis, which reveals that, in the Pv genome, paralogous CNL genes are shared among nonhomologous chromosomes (4 and 11). Together, our results suggest that Pv B4-CNL was derived from CNL sequences from another cluster, the Co-2 cluster, through an ectopic recombination event. Integration of the soybean (Glycine max) genome data enables us to date more precisely this event and also to infer that a single CNL moved from the Co-2 to the B4 cluster. Moreover, we identified a new 528-bp satellite repeat, referred to as khipu, specific to the Phaseolus genus, present both between B4-CNL sequences and in the two knobs identified at the B4 R gene cluster. The khipu repeat is present on most chromosomal termini, indicating the existence of frequent ectopic recombination events in Pv subtelomeric regions. Our results highlight the importance of ectopic recombination in R gene evolution. 2009-11-03 2017-08-28T12:46:03Z 2017-08-28T12:46:03Z Journal Article https://hdl.handle.net/10568/83269 en Limited Access application/pdf Oxford University Press David, Perrine; Chen, Nicolas W.G.; Pedrosa Harand, Andrea; Thareau, Vincent; Sévignac, Mireille; Cannon, Steven B.; Debouck, Daniel G.; Langin, Thierry; Geffroy, Valerie. 2009. A nomadic subtelomeric disease resistance gene cluster in common bean. Plant Physiology 151:1048-1065.
spellingShingle phaseolus vulgaris
phylogeny
recombination
nucleotide sequence
chromosomes
disease resistance
filogenia
recombinación
secuencia nucleotidica
cromosomas
resistencia a la enfermedad
David, Perrine
Chen, Nicolas W.G.
Pedrosa Harand, Andrea
Thareau, Vincent
Sévignac, Mireille
Cannon, Steven B.
Debouck, Daniel G.
Langin, Thierry
Geffroy, Valerie
A nomadic subtelomeric disease resistance gene cluster in common bean
title A nomadic subtelomeric disease resistance gene cluster in common bean
title_full A nomadic subtelomeric disease resistance gene cluster in common bean
title_fullStr A nomadic subtelomeric disease resistance gene cluster in common bean
title_full_unstemmed A nomadic subtelomeric disease resistance gene cluster in common bean
title_short A nomadic subtelomeric disease resistance gene cluster in common bean
title_sort nomadic subtelomeric disease resistance gene cluster in common bean
topic phaseolus vulgaris
phylogeny
recombination
nucleotide sequence
chromosomes
disease resistance
filogenia
recombinación
secuencia nucleotidica
cromosomas
resistencia a la enfermedad
url https://hdl.handle.net/10568/83269
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