Viral class 1 RNase III involved in suppression of RNA silencing

Double-stranded RNA (dsRNA)-specific endonucleases belonging to RNase III classes 3 and 2 process dsRNA precursors to small interfering RNA (siRNA) or microRNA, respectively, thereby initiating and amplifying RNA silencing-based antiviral defense and gene regulation in eukaryotic cells. However, we...

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Main Authors: Kreuze, Jan F., Savenkov, E.I., Cuéllar, Wilmer Jose, Xiangdong Li, Valkonen, J.P.T.
Format: Journal Article
Language:Inglés
Published: American Society for Microbiology 2005
Subjects:
Online Access:https://hdl.handle.net/10568/79401
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author Kreuze, Jan F.
Savenkov, E.I.
Cuéllar, Wilmer Jose
Xiangdong Li
Valkonen, J.P.T.
author_browse Cuéllar, Wilmer Jose
Kreuze, Jan F.
Savenkov, E.I.
Valkonen, J.P.T.
Xiangdong Li
author_facet Kreuze, Jan F.
Savenkov, E.I.
Cuéllar, Wilmer Jose
Xiangdong Li
Valkonen, J.P.T.
author_sort Kreuze, Jan F.
collection Repository of Agricultural Research Outputs (CGSpace)
description Double-stranded RNA (dsRNA)-specific endonucleases belonging to RNase III classes 3 and 2 process dsRNA precursors to small interfering RNA (siRNA) or microRNA, respectively, thereby initiating and amplifying RNA silencing-based antiviral defense and gene regulation in eukaryotic cells. However, we now provide evidence that a class 1 RNase III is involved in suppression of RNA silencing. The single-stranded RNA genome of sweet potato chlorotic stunt virus (SPCSV) encodes an RNase III (RNase3) homologous to putative class 1 RNase IIIs of unknown function in rice and Arabidopsis. We show that RNase3 has dsRNA-specific endonuclease activity that enhances the RNA-silencing suppression activity of another protein (p22) encoded by SPCSV. RNase3 and p22 coexpression reduced siRNA accumulation more efficiently than p22 alone in Nicotiana benthamiana leaves expressing a strong silencing inducer (i.e., dsRNA). RNase3 did not cause intracellular silencing suppression or reduce accumulation of siRNA in the absence of p22 or enhance silencing suppression activity of a protein encoded by a heterologous virus. No other known RNA virus encodes an RNase III or uses two independent proteins cooperatively for RNA silencing suppression.
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spelling CGSpace794012024-04-25T06:00:18Z Viral class 1 RNase III involved in suppression of RNA silencing Kreuze, Jan F. Savenkov, E.I. Cuéllar, Wilmer Jose Xiangdong Li Valkonen, J.P.T. viruses research virology immunology microbiology Double-stranded RNA (dsRNA)-specific endonucleases belonging to RNase III classes 3 and 2 process dsRNA precursors to small interfering RNA (siRNA) or microRNA, respectively, thereby initiating and amplifying RNA silencing-based antiviral defense and gene regulation in eukaryotic cells. However, we now provide evidence that a class 1 RNase III is involved in suppression of RNA silencing. The single-stranded RNA genome of sweet potato chlorotic stunt virus (SPCSV) encodes an RNase III (RNase3) homologous to putative class 1 RNase IIIs of unknown function in rice and Arabidopsis. We show that RNase3 has dsRNA-specific endonuclease activity that enhances the RNA-silencing suppression activity of another protein (p22) encoded by SPCSV. RNase3 and p22 coexpression reduced siRNA accumulation more efficiently than p22 alone in Nicotiana benthamiana leaves expressing a strong silencing inducer (i.e., dsRNA). RNase3 did not cause intracellular silencing suppression or reduce accumulation of siRNA in the absence of p22 or enhance silencing suppression activity of a protein encoded by a heterologous virus. No other known RNA virus encodes an RNase III or uses two independent proteins cooperatively for RNA silencing suppression. 2005-06 2017-01-26T14:21:09Z 2017-01-26T14:21:09Z Journal Article https://hdl.handle.net/10568/79401 en Limited Access American Society for Microbiology Kreuze, J.F.; Savenkov, E.I.; Cuellar, W.; Li, X.; Valkonen, J.P.T. 2005. Viral class 1 RNase III involved in suppression of RNA silencing. Journal of Virology. (USA). ISSN 0022-538X. 79(11):7227-7238.
spellingShingle viruses
research
virology
immunology
microbiology
Kreuze, Jan F.
Savenkov, E.I.
Cuéllar, Wilmer Jose
Xiangdong Li
Valkonen, J.P.T.
Viral class 1 RNase III involved in suppression of RNA silencing
title Viral class 1 RNase III involved in suppression of RNA silencing
title_full Viral class 1 RNase III involved in suppression of RNA silencing
title_fullStr Viral class 1 RNase III involved in suppression of RNA silencing
title_full_unstemmed Viral class 1 RNase III involved in suppression of RNA silencing
title_short Viral class 1 RNase III involved in suppression of RNA silencing
title_sort viral class 1 rnase iii involved in suppression of rna silencing
topic viruses
research
virology
immunology
microbiology
url https://hdl.handle.net/10568/79401
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