IFN-gamma-induced IDO and WRS expression in microglia is differentially regulated by IL-4

Indoleamine 2,3-dioxygenase (IDO), a tryptophan catabolizing enzyme, has been implicated in the pathogenesis of various neurological disorders. IDO expression is induced by IFN-gamma and leads to neurotoxicity by generating quinolinic acid. Additionally, it inhibits the immune response through both...

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Autores principales: Yadav, M.C., Burudi, E.M.E., Alirezaei, M., Flynn, C.C., Watry, D.D., Lanigan, C.M., Fox, H.S.
Formato: Journal Article
Lenguaje:Inglés
Publicado: Wiley 2007
Materias:
Acceso en línea:https://hdl.handle.net/10568/1864
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author Yadav, M.C.
Burudi, E.M.E.
Alirezaei, M.
Flynn, C.C.
Watry, D.D.
Lanigan, C.M.
Fox, H.S.
author_browse Alirezaei, M.
Burudi, E.M.E.
Flynn, C.C.
Fox, H.S.
Lanigan, C.M.
Watry, D.D.
Yadav, M.C.
author_facet Yadav, M.C.
Burudi, E.M.E.
Alirezaei, M.
Flynn, C.C.
Watry, D.D.
Lanigan, C.M.
Fox, H.S.
author_sort Yadav, M.C.
collection Repository of Agricultural Research Outputs (CGSpace)
description Indoleamine 2,3-dioxygenase (IDO), a tryptophan catabolizing enzyme, has been implicated in the pathogenesis of various neurological disorders. IDO expression is induced by IFN-gamma and leads to neurotoxicity by generating quinolinic acid. Additionally, it inhibits the immune response through both tryptophan depletion and generating other tryptophan catabolites. IL-4 and IL-13 have been shown to control IDO expression by antagonizing the effects of IFN-gamma in different cell types. Here, we investigated the effects of these cytokines on IDO expression in microglia. Interestingly, we observed that both IL-4 and IL-13 greatly enhanced IFN-gamma-induced IDO expression. However, tryptophanyl-tRNA synthetase (WRS), which is coinduced with IDO by IFN-gamma, is downregulated by IL-4 and IL-13. The effect of IL-4 and IL-13 was independent of STAT-6. Modulation of IDO but not WRS was eliminated by inhibition of protein phosphatase 2A (PP2A) activity. The phosphatidylinositol 3-kinase (PI3K) pathway further differentiated the regulation of these two enzymes, as inhibiting the PI3K pathway eliminated IFN-gamma induction of IDO, whereas such inhibition greatly enhanced WRS expression. These findings show discordance between modulations of expression of two distinct enzymes utilizing tryptophan as a common substrate, and raise the possibility of their involvement in regulating immune responses in various neurological disorders.
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spelling CGSpace18642023-09-09T12:45:33Z IFN-gamma-induced IDO and WRS expression in microglia is differentially regulated by IL-4 Yadav, M.C. Burudi, E.M.E. Alirezaei, M. Flynn, C.C. Watry, D.D. Lanigan, C.M. Fox, H.S. genetics animal diseases Indoleamine 2,3-dioxygenase (IDO), a tryptophan catabolizing enzyme, has been implicated in the pathogenesis of various neurological disorders. IDO expression is induced by IFN-gamma and leads to neurotoxicity by generating quinolinic acid. Additionally, it inhibits the immune response through both tryptophan depletion and generating other tryptophan catabolites. IL-4 and IL-13 have been shown to control IDO expression by antagonizing the effects of IFN-gamma in different cell types. Here, we investigated the effects of these cytokines on IDO expression in microglia. Interestingly, we observed that both IL-4 and IL-13 greatly enhanced IFN-gamma-induced IDO expression. However, tryptophanyl-tRNA synthetase (WRS), which is coinduced with IDO by IFN-gamma, is downregulated by IL-4 and IL-13. The effect of IL-4 and IL-13 was independent of STAT-6. Modulation of IDO but not WRS was eliminated by inhibition of protein phosphatase 2A (PP2A) activity. The phosphatidylinositol 3-kinase (PI3K) pathway further differentiated the regulation of these two enzymes, as inhibiting the PI3K pathway eliminated IFN-gamma induction of IDO, whereas such inhibition greatly enhanced WRS expression. These findings show discordance between modulations of expression of two distinct enzymes utilizing tryptophan as a common substrate, and raise the possibility of their involvement in regulating immune responses in various neurological disorders. 2007-10 2010-06-13T16:00:50Z 2010-06-13T16:00:50Z Journal Article https://hdl.handle.net/10568/1864 en Limited Access Wiley Yadav, M.C.; Burudi, E.M.; Alirezaei, M.; Flynn, C.C.; Watry, D.D.; Lanigan, C.M., Fox, H.S. 2007. IFN-gamma-induced IDO and WRS expression in microglia is differentially regulated by IL-4. Glia 55(13):1385-1396.
spellingShingle genetics
animal diseases
Yadav, M.C.
Burudi, E.M.E.
Alirezaei, M.
Flynn, C.C.
Watry, D.D.
Lanigan, C.M.
Fox, H.S.
IFN-gamma-induced IDO and WRS expression in microglia is differentially regulated by IL-4
title IFN-gamma-induced IDO and WRS expression in microglia is differentially regulated by IL-4
title_full IFN-gamma-induced IDO and WRS expression in microglia is differentially regulated by IL-4
title_fullStr IFN-gamma-induced IDO and WRS expression in microglia is differentially regulated by IL-4
title_full_unstemmed IFN-gamma-induced IDO and WRS expression in microglia is differentially regulated by IL-4
title_short IFN-gamma-induced IDO and WRS expression in microglia is differentially regulated by IL-4
title_sort ifn gamma induced ido and wrs expression in microglia is differentially regulated by il 4
topic genetics
animal diseases
url https://hdl.handle.net/10568/1864
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