Revisiting the relationship between stomatal size and speed across species – a meta‐analysis

The rate of stomatal opening and closure in response to changes in light affects leaf photosynthesis and water use. However, it is unclear how strongly stomatal size ( SS ) and density ( SD ) influence stomatal conductance ( g s ) kinetics, and whether variation arises from methodological difference...

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Autores principales: Woning, Nik, Al‐Salman, Yazen, Kaiser, Elias, Berman, Sarah R., Brendel, Oliver, Cano, Francisco Javier, Carpentier, Sebastien, Centritto, Mauro, Drake, Paul L., Durand, Maxime, Eyland, David, Franks, Peter J., Gerardin, Theo, Ghannoum, Oula, Haworth, Matthew, Kübarsepp, Liisa, Lawson, Tracy, Le Thiec, Didier, Li, Yong, Marcelis, Leo F. M., Marino, Giovanni, McAusland, Lorna, Muir, Christopher D., Niinemets, Ülo, Nunes, Tiago D. G., Raissig, Michael T., Sakoda, Kazuma, Sugiura, Daisuke, Tosens, Tiina, Zhang, Qiangqiang, Zhang, Ningyi, Vialet‐Chabrand, Silvere
Formato: Journal Article
Lenguaje:Inglés
Publicado: Wiley 2025
Materias:
Acceso en línea:https://hdl.handle.net/10568/179478
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author Woning, Nik
Al‐Salman, Yazen
Kaiser, Elias
Berman, Sarah R.
Brendel, Oliver
Cano, Francisco Javier
Carpentier, Sebastien
Centritto, Mauro
Drake, Paul L.
Durand, Maxime
Eyland, David
Franks, Peter J.
Gerardin, Theo
Ghannoum, Oula
Haworth, Matthew
Kübarsepp, Liisa
Lawson, Tracy
Le Thiec, Didier
Li, Yong
Marcelis, Leo F. M.
Marino, Giovanni
McAusland, Lorna
Muir, Christopher D.
Niinemets, Ülo
Nunes, Tiago D. G.
Raissig, Michael T.
Sakoda, Kazuma
Sugiura, Daisuke
Tosens, Tiina
Zhang, Qiangqiang
Zhang, Ningyi
Vialet‐Chabrand, Silvere
author_browse Al‐Salman, Yazen
Berman, Sarah R.
Brendel, Oliver
Cano, Francisco Javier
Carpentier, Sebastien
Centritto, Mauro
Drake, Paul L.
Durand, Maxime
Eyland, David
Franks, Peter J.
Gerardin, Theo
Ghannoum, Oula
Haworth, Matthew
Kaiser, Elias
Kübarsepp, Liisa
Lawson, Tracy
Le Thiec, Didier
Li, Yong
Marcelis, Leo F. M.
Marino, Giovanni
McAusland, Lorna
Muir, Christopher D.
Niinemets, Ülo
Nunes, Tiago D. G.
Raissig, Michael T.
Sakoda, Kazuma
Sugiura, Daisuke
Tosens, Tiina
Vialet‐Chabrand, Silvere
Woning, Nik
Zhang, Ningyi
Zhang, Qiangqiang
author_facet Woning, Nik
Al‐Salman, Yazen
Kaiser, Elias
Berman, Sarah R.
Brendel, Oliver
Cano, Francisco Javier
Carpentier, Sebastien
Centritto, Mauro
Drake, Paul L.
Durand, Maxime
Eyland, David
Franks, Peter J.
Gerardin, Theo
Ghannoum, Oula
Haworth, Matthew
Kübarsepp, Liisa
Lawson, Tracy
Le Thiec, Didier
Li, Yong
Marcelis, Leo F. M.
Marino, Giovanni
McAusland, Lorna
Muir, Christopher D.
Niinemets, Ülo
Nunes, Tiago D. G.
Raissig, Michael T.
Sakoda, Kazuma
Sugiura, Daisuke
Tosens, Tiina
Zhang, Qiangqiang
Zhang, Ningyi
Vialet‐Chabrand, Silvere
author_sort Woning, Nik
collection Repository of Agricultural Research Outputs (CGSpace)
description The rate of stomatal opening and closure in response to changes in light affects leaf photosynthesis and water use. However, it is unclear how strongly stomatal size ( SS ) and density ( SD ) influence stomatal conductance ( g s ) kinetics, and whether variation arises from methodological differences, guard cell type or degree of amphistomaty. We divided published records combining stomatal kinetics and anatomical traits from 89 species into kidney and dumbbell‐shaped guard cells, and evaluated four dynamic g s models on them. We derived the time constant for an exponential response of g s ( τ ) and the maximum rate of change ( Sl max ) as well as the ratio of adaxial/abaxial SD ( rSD ). We found significant differences in parameter estimation between models. Stomatal anatomical traits and kinetic parameters showed large variation across species. While individual anatomical features ( SS , SD , rSD and guard cell types) were weakly correlated with stomatal response speed ( τ and Sl max ), interactions between these features showed significant effects, demonstrating that kinetic performance arises from synergistic rather than additive anatomical relationships. Our results call for the use of our unified modeling approach, challenge the generality of the observation that smaller stomata move faster across species and suggest rSD as an understudied driver of stomatal kinetics.
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spelling CGSpace1794782026-01-08T02:07:23Z Revisiting the relationship between stomatal size and speed across species – a meta‐analysis Woning, Nik Al‐Salman, Yazen Kaiser, Elias Berman, Sarah R. Brendel, Oliver Cano, Francisco Javier Carpentier, Sebastien Centritto, Mauro Drake, Paul L. Durand, Maxime Eyland, David Franks, Peter J. Gerardin, Theo Ghannoum, Oula Haworth, Matthew Kübarsepp, Liisa Lawson, Tracy Le Thiec, Didier Li, Yong Marcelis, Leo F. M. Marino, Giovanni McAusland, Lorna Muir, Christopher D. Niinemets, Ülo Nunes, Tiago D. G. Raissig, Michael T. Sakoda, Kazuma Sugiura, Daisuke Tosens, Tiina Zhang, Qiangqiang Zhang, Ningyi Vialet‐Chabrand, Silvere stomatal conductance modelling species diversity kinetics The rate of stomatal opening and closure in response to changes in light affects leaf photosynthesis and water use. However, it is unclear how strongly stomatal size ( SS ) and density ( SD ) influence stomatal conductance ( g s ) kinetics, and whether variation arises from methodological differences, guard cell type or degree of amphistomaty. We divided published records combining stomatal kinetics and anatomical traits from 89 species into kidney and dumbbell‐shaped guard cells, and evaluated four dynamic g s models on them. We derived the time constant for an exponential response of g s ( τ ) and the maximum rate of change ( Sl max ) as well as the ratio of adaxial/abaxial SD ( rSD ). We found significant differences in parameter estimation between models. Stomatal anatomical traits and kinetic parameters showed large variation across species. While individual anatomical features ( SS , SD , rSD and guard cell types) were weakly correlated with stomatal response speed ( τ and Sl max ), interactions between these features showed significant effects, demonstrating that kinetic performance arises from synergistic rather than additive anatomical relationships. Our results call for the use of our unified modeling approach, challenge the generality of the observation that smaller stomata move faster across species and suggest rSD as an understudied driver of stomatal kinetics. 2025-12-28 2026-01-07T12:59:53Z 2026-01-07T12:59:53Z Journal Article https://hdl.handle.net/10568/179478 en Open Access application/pdf Wiley Woning, N.; Al‐Salman, Y.; Kaiser, E.; Berman, S.R.; Brendel, O.; Cano, F.J.; Carpentier, S.; Centritto, M.; Drake, P.L.; Durand, M.; Eyland, D.; Franks, P.J.; Gerardin, T.; Ghannoum, O.; Haworth, M.; Kübarsepp, L.; Lawson, T.; Le Thiec, D.; Li, Y.; Marcelis, L.F.M.; Marino, G.; McAusland, L.; Muir, C.D.; Niinemets, Ü.; Nunes, T.D.G.; Raissig, M.T.; Sakoda, K.; Sugiura, D.; Tosens, T.; Zhang, Q.; Zhang, N.; Vialet‐Chabrand, S. (2025) Revisiting the relationship between stomatal size and speed across species – a meta‐analysis. New Phytologist , Online first paper(2025-12-28). ISSN: 0028-646X
spellingShingle stomatal conductance
modelling
species diversity
kinetics
Woning, Nik
Al‐Salman, Yazen
Kaiser, Elias
Berman, Sarah R.
Brendel, Oliver
Cano, Francisco Javier
Carpentier, Sebastien
Centritto, Mauro
Drake, Paul L.
Durand, Maxime
Eyland, David
Franks, Peter J.
Gerardin, Theo
Ghannoum, Oula
Haworth, Matthew
Kübarsepp, Liisa
Lawson, Tracy
Le Thiec, Didier
Li, Yong
Marcelis, Leo F. M.
Marino, Giovanni
McAusland, Lorna
Muir, Christopher D.
Niinemets, Ülo
Nunes, Tiago D. G.
Raissig, Michael T.
Sakoda, Kazuma
Sugiura, Daisuke
Tosens, Tiina
Zhang, Qiangqiang
Zhang, Ningyi
Vialet‐Chabrand, Silvere
Revisiting the relationship between stomatal size and speed across species – a meta‐analysis
title Revisiting the relationship between stomatal size and speed across species – a meta‐analysis
title_full Revisiting the relationship between stomatal size and speed across species – a meta‐analysis
title_fullStr Revisiting the relationship between stomatal size and speed across species – a meta‐analysis
title_full_unstemmed Revisiting the relationship between stomatal size and speed across species – a meta‐analysis
title_short Revisiting the relationship between stomatal size and speed across species – a meta‐analysis
title_sort revisiting the relationship between stomatal size and speed across species a meta analysis
topic stomatal conductance
modelling
species diversity
kinetics
url https://hdl.handle.net/10568/179478
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