Bundle sheath suberisation is required for C4 photosynthesis in a Setaria viridis mutant

C4 photosynthesis provides an effective solution for overcoming the catalytic inefficiency of Rubisco. The pathway is characterised by a biochemical CO2 concentrating mechanism that operates across mesophyll and bundle sheath (BS) cells and relies on a gas tight BS compartment. A screen of a mutant...

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Main Authors: Danila, Florence R., Thakur, Vivek, Chatterjee, Jolly, Bala, Soumi, Coe, Robert A., Acebron, Kelvin, Furbank, Robert T., von Caemmerer, Susanne, Quick, William Paul
Format: Journal Article
Language:Inglés
Published: Springer 2021
Online Access:https://hdl.handle.net/10568/164328
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author Danila, Florence R.
Thakur, Vivek
Chatterjee, Jolly
Bala, Soumi
Coe, Robert A.
Acebron, Kelvin
Furbank, Robert T.
von Caemmerer, Susanne
Quick, William Paul
author_browse Acebron, Kelvin
Bala, Soumi
Chatterjee, Jolly
Coe, Robert A.
Danila, Florence R.
Furbank, Robert T.
Quick, William Paul
Thakur, Vivek
von Caemmerer, Susanne
author_facet Danila, Florence R.
Thakur, Vivek
Chatterjee, Jolly
Bala, Soumi
Coe, Robert A.
Acebron, Kelvin
Furbank, Robert T.
von Caemmerer, Susanne
Quick, William Paul
author_sort Danila, Florence R.
collection Repository of Agricultural Research Outputs (CGSpace)
description C4 photosynthesis provides an effective solution for overcoming the catalytic inefficiency of Rubisco. The pathway is characterised by a biochemical CO2 concentrating mechanism that operates across mesophyll and bundle sheath (BS) cells and relies on a gas tight BS compartment. A screen of a mutant population of Setaria viridis, an NADP-malic enzyme type C4 monocot, generated using N-nitroso-N-methylurea identified a mutant with an amino acid change in the gene coding region of the ABCG transporter, a step in the suberin synthesis pathway. Here, Nile red staining, TEM, and GC/MS confirmed the alteration in suberin deposition in the BS cell wall of the mutant. We show that this has disrupted the suberin lamellae of BS cell wall and increased BS conductance to CO2 diffusion more than two-fold in the mutant. Consequently, BS CO2 partial pressure is reduced and CO2 assimilation was impaired in the mutant. Our findings provide experimental evidence that a functional suberin lamellae is an essential anatomical feature for efficient C4 photosynthesis in NADP-ME plants like S. viridis and have implications for engineering strategies to ensure future food security.
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spelling CGSpace1643282024-12-19T14:12:02Z Bundle sheath suberisation is required for C4 photosynthesis in a Setaria viridis mutant Danila, Florence R. Thakur, Vivek Chatterjee, Jolly Bala, Soumi Coe, Robert A. Acebron, Kelvin Furbank, Robert T. von Caemmerer, Susanne Quick, William Paul C4 photosynthesis provides an effective solution for overcoming the catalytic inefficiency of Rubisco. The pathway is characterised by a biochemical CO2 concentrating mechanism that operates across mesophyll and bundle sheath (BS) cells and relies on a gas tight BS compartment. A screen of a mutant population of Setaria viridis, an NADP-malic enzyme type C4 monocot, generated using N-nitroso-N-methylurea identified a mutant with an amino acid change in the gene coding region of the ABCG transporter, a step in the suberin synthesis pathway. Here, Nile red staining, TEM, and GC/MS confirmed the alteration in suberin deposition in the BS cell wall of the mutant. We show that this has disrupted the suberin lamellae of BS cell wall and increased BS conductance to CO2 diffusion more than two-fold in the mutant. Consequently, BS CO2 partial pressure is reduced and CO2 assimilation was impaired in the mutant. Our findings provide experimental evidence that a functional suberin lamellae is an essential anatomical feature for efficient C4 photosynthesis in NADP-ME plants like S. viridis and have implications for engineering strategies to ensure future food security. 2021-02-26 2024-12-19T12:53:44Z 2024-12-19T12:53:44Z Journal Article https://hdl.handle.net/10568/164328 en Open Access Springer Danila, Florence R.; Thakur, Vivek; Chatterjee, Jolly; Bala, Soumi; Coe, Robert A.; Acebron, Kelvin; Furbank, Robert T.; von Caemmerer, Susanne and Quick, William Paul. 2021. Bundle sheath suberisation is required for C4 photosynthesis in a Setaria viridis mutant. Commun Biol, Volume 4, no. 1
spellingShingle Danila, Florence R.
Thakur, Vivek
Chatterjee, Jolly
Bala, Soumi
Coe, Robert A.
Acebron, Kelvin
Furbank, Robert T.
von Caemmerer, Susanne
Quick, William Paul
Bundle sheath suberisation is required for C4 photosynthesis in a Setaria viridis mutant
title Bundle sheath suberisation is required for C4 photosynthesis in a Setaria viridis mutant
title_full Bundle sheath suberisation is required for C4 photosynthesis in a Setaria viridis mutant
title_fullStr Bundle sheath suberisation is required for C4 photosynthesis in a Setaria viridis mutant
title_full_unstemmed Bundle sheath suberisation is required for C4 photosynthesis in a Setaria viridis mutant
title_short Bundle sheath suberisation is required for C4 photosynthesis in a Setaria viridis mutant
title_sort bundle sheath suberisation is required for c4 photosynthesis in a setaria viridis mutant
url https://hdl.handle.net/10568/164328
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