Nitrogen form-mediated ethylene signal regulates root-to-shoot K+ translocation via NRT1.5

Nitrogen–potassium synergistic and antagonistic interactions are the typical case of nutrient interactions. However, the underlying mechanism for the integration of the external N form into K+ homeostasis remains unclear. Here, we show that opposite effects of NO3− and NH4+ on root‐shoot K+ transloc...

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Autores principales: Chen, Haifei, Zhang, Quan, Wang, Xueru, Zhang, Jianhua, Ismail, Abdelbagi M., Zhang, Zhenhua
Formato: Journal Article
Lenguaje:Inglés
Publicado: Wiley 2021
Materias:
Acceso en línea:https://hdl.handle.net/10568/164192
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author Chen, Haifei
Zhang, Quan
Wang, Xueru
Zhang, Jianhua
Ismail, Abdelbagi M.
Zhang, Zhenhua
author_browse Chen, Haifei
Ismail, Abdelbagi M.
Wang, Xueru
Zhang, Jianhua
Zhang, Quan
Zhang, Zhenhua
author_facet Chen, Haifei
Zhang, Quan
Wang, Xueru
Zhang, Jianhua
Ismail, Abdelbagi M.
Zhang, Zhenhua
author_sort Chen, Haifei
collection Repository of Agricultural Research Outputs (CGSpace)
description Nitrogen–potassium synergistic and antagonistic interactions are the typical case of nutrient interactions. However, the underlying mechanism for the integration of the external N form into K+ homeostasis remains unclear. Here, we show that opposite effects of NO3− and NH4+ on root‐shoot K+ translocation were due to differential regulation of an ethylene signalling pathway targeting the NRT1.5 transporter. NH4+ upregulated the transcriptional activity of EIN3, but repressed the expression of NRT1.5. However, the addition of NO3− strongly suppressed the activity of EIN3, whereas its addition upregulated the expression of AtNRT1.5 and shoot K+ concentration. The 35S:EIN3/ein3eil1 plants, nrt1.5 mutants and nrt1.5/skor double mutants displayed a low K+ chlorosis phenotype, especially under NH4+ conditions with low K+ supply. Ion content analyses indicate that root‐to‐shoot K+ translocation was significantly reduced in these mutants. A Y1H assay, an EMSA and a transient expression assay confirmed that AtEIN3 protein could directly bind to the promoter of NRT1.5 to repress its expression. Furthermore, grafted plants with the roots of 35S:EIN3 and ein3eil1/nrt1.5 mutants displayed marked leaf chlorosis with a low K+ concentration. Collectively, our findings reveal that the interaction between N form and K+ was achieved by modulating root‐derived ethylene signals to regulate root‐to‐shoot K+ translocation via NRT1.5.
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spelling CGSpace1641922025-01-24T14:20:08Z Nitrogen form-mediated ethylene signal regulates root-to-shoot K+ translocation via NRT1.5 Chen, Haifei Zhang, Quan Wang, Xueru Zhang, Jianhua Ismail, Abdelbagi M. Zhang, Zhenhua plant science physiology Nitrogen–potassium synergistic and antagonistic interactions are the typical case of nutrient interactions. However, the underlying mechanism for the integration of the external N form into K+ homeostasis remains unclear. Here, we show that opposite effects of NO3− and NH4+ on root‐shoot K+ translocation were due to differential regulation of an ethylene signalling pathway targeting the NRT1.5 transporter. NH4+ upregulated the transcriptional activity of EIN3, but repressed the expression of NRT1.5. However, the addition of NO3− strongly suppressed the activity of EIN3, whereas its addition upregulated the expression of AtNRT1.5 and shoot K+ concentration. The 35S:EIN3/ein3eil1 plants, nrt1.5 mutants and nrt1.5/skor double mutants displayed a low K+ chlorosis phenotype, especially under NH4+ conditions with low K+ supply. Ion content analyses indicate that root‐to‐shoot K+ translocation was significantly reduced in these mutants. A Y1H assay, an EMSA and a transient expression assay confirmed that AtEIN3 protein could directly bind to the promoter of NRT1.5 to repress its expression. Furthermore, grafted plants with the roots of 35S:EIN3 and ein3eil1/nrt1.5 mutants displayed marked leaf chlorosis with a low K+ concentration. Collectively, our findings reveal that the interaction between N form and K+ was achieved by modulating root‐derived ethylene signals to regulate root‐to‐shoot K+ translocation via NRT1.5. 2021-12 2024-12-19T12:53:35Z 2024-12-19T12:53:35Z Journal Article https://hdl.handle.net/10568/164192 en Wiley Chen, Haifei; Zhang, Quan; Wang, Xueru; Zhang, Jianhua; Ismail, Abdelbagi M. and Zhang, Zhenhua. 2021. Nitrogen form-mediated ethylene signal regulates root-to-shoot K+ translocation via NRT1.5. Plant Cell and Environment, Volume 44 no. 12 p. 3806-3818
spellingShingle plant science
physiology
Chen, Haifei
Zhang, Quan
Wang, Xueru
Zhang, Jianhua
Ismail, Abdelbagi M.
Zhang, Zhenhua
Nitrogen form-mediated ethylene signal regulates root-to-shoot K+ translocation via NRT1.5
title Nitrogen form-mediated ethylene signal regulates root-to-shoot K+ translocation via NRT1.5
title_full Nitrogen form-mediated ethylene signal regulates root-to-shoot K+ translocation via NRT1.5
title_fullStr Nitrogen form-mediated ethylene signal regulates root-to-shoot K+ translocation via NRT1.5
title_full_unstemmed Nitrogen form-mediated ethylene signal regulates root-to-shoot K+ translocation via NRT1.5
title_short Nitrogen form-mediated ethylene signal regulates root-to-shoot K+ translocation via NRT1.5
title_sort nitrogen form mediated ethylene signal regulates root to shoot k translocation via nrt1 5
topic plant science
physiology
url https://hdl.handle.net/10568/164192
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