Gene B-5 in cotton confers high and broad resistance to bacterial blight and conditions high amounts of sesquiterpenoid phytoalexins

Bacterial blight resistance gene B5 has received little attention since it was first described in 1950. A near-isogenic line (NIL) of Gossypium hirsutum cotton, Ac B5, was generated in an otherwise bacterial-blight-susceptible ‘Acala 44’ background. The introgressed locus B5 in Ac B5 conferred stron...

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Main Authors: Essenberg, Margaret, McNally, Kenneth L., Bayles, Melanie B., Pierce, Margaret L., Hall, Judy A., Kuss, Christine R., Shevell, Judith L., Verhalen, Laval M.
Format: Journal Article
Language:Inglés
Published: Scientific Societies 2023
Subjects:
Online Access:https://hdl.handle.net/10568/163956
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author Essenberg, Margaret
McNally, Kenneth L.
Bayles, Melanie B.
Pierce, Margaret L.
Hall, Judy A.
Kuss, Christine R.
Shevell, Judith L.
Verhalen, Laval M.
author_browse Bayles, Melanie B.
Essenberg, Margaret
Hall, Judy A.
Kuss, Christine R.
McNally, Kenneth L.
Pierce, Margaret L.
Shevell, Judith L.
Verhalen, Laval M.
author_facet Essenberg, Margaret
McNally, Kenneth L.
Bayles, Melanie B.
Pierce, Margaret L.
Hall, Judy A.
Kuss, Christine R.
Shevell, Judith L.
Verhalen, Laval M.
author_sort Essenberg, Margaret
collection Repository of Agricultural Research Outputs (CGSpace)
description Bacterial blight resistance gene B5 has received little attention since it was first described in 1950. A near-isogenic line (NIL) of Gossypium hirsutum cotton, Ac B5, was generated in an otherwise bacterial-blight-susceptible ‘Acala 44’ background. The introgressed locus B5 in Ac B5 conferred strong and broad-spectrum resistance to bacterial blight. Segregation patterns of test crosses under Oklahoma field conditions indicated that Ac B5 is likely homozygous for resistance at two loci with partial dominance gene action. In controlled-environment conditions, two of the four copies of B5 were required for effective resistance. Contrary to expectations of gene-for-gene theory, Ac B5 conferred high resistance toward isogenic strains of Xanthomonas citri subsp. malvacearum carrying cloned avirulence genes avrB4, avrb7, avrBIn, avrB101, and avrB102, respectively, and weaker resistance toward the strain carrying cloned avrb6. The hypothesis that each B gene, in the absence of a polygenic complex, triggers sesquiterpenoid phytoalexin production was tested by measurement of cadalene and lacinilene phytoalexins during resistant responses in five NILs carrying different B genes, four other lines carrying multiple resistance genes, as well as susceptible Ac44E. Phytoalexin production was an obvious, but variable, response in all nine resistant lines. Ac B5 accumulated an order of magnitude more of all four phytoalexins than any of the other resistant NILs. Its total levels were comparable to those detected in OK1.2, a highly resistant line that possesses several B genes in a polygenic background.
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spelling CGSpace1639562024-12-22T05:45:03Z Gene B-5 in cotton confers high and broad resistance to bacterial blight and conditions high amounts of sesquiterpenoid phytoalexins Essenberg, Margaret McNally, Kenneth L. Bayles, Melanie B. Pierce, Margaret L. Hall, Judy A. Kuss, Christine R. Shevell, Judith L. Verhalen, Laval M. genes disease resistance phytoalexins gossypium Bacterial blight resistance gene B5 has received little attention since it was first described in 1950. A near-isogenic line (NIL) of Gossypium hirsutum cotton, Ac B5, was generated in an otherwise bacterial-blight-susceptible ‘Acala 44’ background. The introgressed locus B5 in Ac B5 conferred strong and broad-spectrum resistance to bacterial blight. Segregation patterns of test crosses under Oklahoma field conditions indicated that Ac B5 is likely homozygous for resistance at two loci with partial dominance gene action. In controlled-environment conditions, two of the four copies of B5 were required for effective resistance. Contrary to expectations of gene-for-gene theory, Ac B5 conferred high resistance toward isogenic strains of Xanthomonas citri subsp. malvacearum carrying cloned avirulence genes avrB4, avrb7, avrBIn, avrB101, and avrB102, respectively, and weaker resistance toward the strain carrying cloned avrb6. The hypothesis that each B gene, in the absence of a polygenic complex, triggers sesquiterpenoid phytoalexin production was tested by measurement of cadalene and lacinilene phytoalexins during resistant responses in five NILs carrying different B genes, four other lines carrying multiple resistance genes, as well as susceptible Ac44E. Phytoalexin production was an obvious, but variable, response in all nine resistant lines. Ac B5 accumulated an order of magnitude more of all four phytoalexins than any of the other resistant NILs. Its total levels were comparable to those detected in OK1.2, a highly resistant line that possesses several B genes in a polygenic background. 2023-05 2024-12-19T12:53:13Z 2024-12-19T12:53:13Z Journal Article https://hdl.handle.net/10568/163956 en Scientific Societies Essenberg, M., McNally, K. L., Bayles, M. B., Pierce, M. L., Hall, J. A., Kuss, C. R., Shevell, J. L., & Verhalen, L. M. (2023). Gene B5 in Cotton Confers High and Broad Resistance to Bacterial Blight and Conditions High Amounts of Sesquiterpenoid Phytoalexins. Phytopathology®, 113(5), 812-823. https://doi.org/10.1094/phyto-08-22-0310-fi
spellingShingle genes
disease resistance
phytoalexins
gossypium
Essenberg, Margaret
McNally, Kenneth L.
Bayles, Melanie B.
Pierce, Margaret L.
Hall, Judy A.
Kuss, Christine R.
Shevell, Judith L.
Verhalen, Laval M.
Gene B-5 in cotton confers high and broad resistance to bacterial blight and conditions high amounts of sesquiterpenoid phytoalexins
title Gene B-5 in cotton confers high and broad resistance to bacterial blight and conditions high amounts of sesquiterpenoid phytoalexins
title_full Gene B-5 in cotton confers high and broad resistance to bacterial blight and conditions high amounts of sesquiterpenoid phytoalexins
title_fullStr Gene B-5 in cotton confers high and broad resistance to bacterial blight and conditions high amounts of sesquiterpenoid phytoalexins
title_full_unstemmed Gene B-5 in cotton confers high and broad resistance to bacterial blight and conditions high amounts of sesquiterpenoid phytoalexins
title_short Gene B-5 in cotton confers high and broad resistance to bacterial blight and conditions high amounts of sesquiterpenoid phytoalexins
title_sort gene b 5 in cotton confers high and broad resistance to bacterial blight and conditions high amounts of sesquiterpenoid phytoalexins
topic genes
disease resistance
phytoalexins
gossypium
url https://hdl.handle.net/10568/163956
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