Natural variation of OsML1, a mitochondrial transcription termination factor, contributes to mesocotyl length variation in rice

Mesocotyl length (ML) is a crucial factor in determining the establishment and yield of rice planted through dry direct seeding, a practice that is increasingly popular in rice production worldwide. ML is determined by the endogenous and external environments, and inherits as a complex trait. To dat...

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Main Authors: Meng, Yun, Zhan, Junhui, Liu, Hongyan, Liu, Jindong, Wang, Yamei, Guo, Zhan, He, Sang, Nie, Lixiao, Kohli, Ajay, Ye, Guoyou
Format: Journal Article
Language:Inglés
Published: Wiley 2023
Subjects:
Online Access:https://hdl.handle.net/10568/163948
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author Meng, Yun
Zhan, Junhui
Liu, Hongyan
Liu, Jindong
Wang, Yamei
Guo, Zhan
He, Sang
Nie, Lixiao
Kohli, Ajay
Ye, Guoyou
author_browse Guo, Zhan
He, Sang
Kohli, Ajay
Liu, Hongyan
Liu, Jindong
Meng, Yun
Nie, Lixiao
Wang, Yamei
Ye, Guoyou
Zhan, Junhui
author_facet Meng, Yun
Zhan, Junhui
Liu, Hongyan
Liu, Jindong
Wang, Yamei
Guo, Zhan
He, Sang
Nie, Lixiao
Kohli, Ajay
Ye, Guoyou
author_sort Meng, Yun
collection Repository of Agricultural Research Outputs (CGSpace)
description Mesocotyl length (ML) is a crucial factor in determining the establishment and yield of rice planted through dry direct seeding, a practice that is increasingly popular in rice production worldwide. ML is determined by the endogenous and external environments, and inherits as a complex trait. To date, only a few genes have been cloned, and the mechanisms underlying mesocotyl elongation remain largely unknown. Here, through a genome‐wide association study using sequenced germplasm, we reveal that natural allelic variations in a mitochondrial transcription termination factor, OsML1, predominantly determined the natural variation of ML in rice. Natural variants in the coding regions of OsML1 resulted in five major haplotypes with a clear differentiation between subspecies and subpopulations in cultivated rice. The much‐reduced genetic diversity of cultivated rice compared to the common wild rice suggested that OsML1 underwent selection during domestication. Transgenic experiments and molecular analysis demonstrated that OsML1 contributes to ML by influencing cell elongation primarily determined by H2O2 homeostasis. Overexpression of OsML1 promoted mesocotyl elongation and thus improved the emergence rate under deep direct seeding. Taken together, our results suggested that OsML1 is a key positive regulator of ML, and is useful in developing varieties for deep direct seeding by conventional and transgenic approaches.
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spelling CGSpace1639482024-12-22T05:45:01Z Natural variation of OsML1, a mitochondrial transcription termination factor, contributes to mesocotyl length variation in rice Meng, Yun Zhan, Junhui Liu, Hongyan Liu, Jindong Wang, Yamei Guo, Zhan He, Sang Nie, Lixiao Kohli, Ajay Ye, Guoyou direct sowing allelic heterogeneity haplotypes genetic variation domestication seedling emergence cell elongation Mesocotyl length (ML) is a crucial factor in determining the establishment and yield of rice planted through dry direct seeding, a practice that is increasingly popular in rice production worldwide. ML is determined by the endogenous and external environments, and inherits as a complex trait. To date, only a few genes have been cloned, and the mechanisms underlying mesocotyl elongation remain largely unknown. Here, through a genome‐wide association study using sequenced germplasm, we reveal that natural allelic variations in a mitochondrial transcription termination factor, OsML1, predominantly determined the natural variation of ML in rice. Natural variants in the coding regions of OsML1 resulted in five major haplotypes with a clear differentiation between subspecies and subpopulations in cultivated rice. The much‐reduced genetic diversity of cultivated rice compared to the common wild rice suggested that OsML1 underwent selection during domestication. Transgenic experiments and molecular analysis demonstrated that OsML1 contributes to ML by influencing cell elongation primarily determined by H2O2 homeostasis. Overexpression of OsML1 promoted mesocotyl elongation and thus improved the emergence rate under deep direct seeding. Taken together, our results suggested that OsML1 is a key positive regulator of ML, and is useful in developing varieties for deep direct seeding by conventional and transgenic approaches. 2023-08 2024-12-19T12:53:13Z 2024-12-19T12:53:13Z Journal Article https://hdl.handle.net/10568/163948 en Wiley Meng, Yun; Zhan, Junhui; Liu, Hongyan; Liu, Jindong; Wang, Yamei; Guo, Zhan; He, Sang; Nie, Lixiao; Kohli, Ajay and Ye, Guoyou. 2023. Natural variation of OsML1, a mitochondrial transcription termination factor, contributes to mesocotyl length variation in rice. The Plant Journal, Volume 115 no. 4 p. 910-925
spellingShingle direct sowing
allelic heterogeneity
haplotypes
genetic variation
domestication
seedling emergence
cell elongation
Meng, Yun
Zhan, Junhui
Liu, Hongyan
Liu, Jindong
Wang, Yamei
Guo, Zhan
He, Sang
Nie, Lixiao
Kohli, Ajay
Ye, Guoyou
Natural variation of OsML1, a mitochondrial transcription termination factor, contributes to mesocotyl length variation in rice
title Natural variation of OsML1, a mitochondrial transcription termination factor, contributes to mesocotyl length variation in rice
title_full Natural variation of OsML1, a mitochondrial transcription termination factor, contributes to mesocotyl length variation in rice
title_fullStr Natural variation of OsML1, a mitochondrial transcription termination factor, contributes to mesocotyl length variation in rice
title_full_unstemmed Natural variation of OsML1, a mitochondrial transcription termination factor, contributes to mesocotyl length variation in rice
title_short Natural variation of OsML1, a mitochondrial transcription termination factor, contributes to mesocotyl length variation in rice
title_sort natural variation of osml1 a mitochondrial transcription termination factor contributes to mesocotyl length variation in rice
topic direct sowing
allelic heterogeneity
haplotypes
genetic variation
domestication
seedling emergence
cell elongation
url https://hdl.handle.net/10568/163948
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