Extensive Polymorphism and Evidence of Immune Selection in a Highly Dominant Antigen Recognized by Bovine CD8 T Cells Specific for Theileria annulata

Although parasite strain-restricted CD8 T cell responses have been described for several protozoa, the precise role of antigenic variability in immunity is poorly understood. The tick-borne protozoan parasiteTheileria annulatainfects leukocytes and causes an acute, often fatal lymphoproliferative di...

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Autores principales: MacHugh, Niall D., Weir, William, Burrells, Alison, Lizundia, Regina, Graham, Simon P., Taracha, Evans L., Shiels, Brian R., Langsley, Gordon, Morrison, W. Ivan
Formato: Journal Article
Lenguaje:Inglés
Publicado: American Society for Microbiology 2011
Materias:
Acceso en línea:https://hdl.handle.net/10568/129596
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author MacHugh, Niall D.
Weir, William
Burrells, Alison
Lizundia, Regina
Graham, Simon P.
Taracha, Evans L.
Shiels, Brian R.
Langsley, Gordon
Morrison, W. Ivan
author_browse Burrells, Alison
Graham, Simon P.
Langsley, Gordon
Lizundia, Regina
MacHugh, Niall D.
Morrison, W. Ivan
Shiels, Brian R.
Taracha, Evans L.
Weir, William
author_facet MacHugh, Niall D.
Weir, William
Burrells, Alison
Lizundia, Regina
Graham, Simon P.
Taracha, Evans L.
Shiels, Brian R.
Langsley, Gordon
Morrison, W. Ivan
author_sort MacHugh, Niall D.
collection Repository of Agricultural Research Outputs (CGSpace)
description Although parasite strain-restricted CD8 T cell responses have been described for several protozoa, the precise role of antigenic variability in immunity is poorly understood. The tick-borne protozoan parasiteTheileria annulatainfects leukocytes and causes an acute, often fatal lymphoproliferative disease in cattle. Building on previous evidence of strain-restricted CD8 T cell responses toT. annulata, this study set out to identify and characterize the variability of the target antigens. Three antigens were identified by screening expressed parasite cDNAs with specific CD8 T cell lines. In cattle expressing the A10 class I major histocompatibility complex haplotype, A10-restricted CD8 T cell responses were shown to be focused entirely on a single dominant epitope in one of these antigens (Ta9). Sequencing of the Ta9 gene from field isolates ofT. annulatademonstrated extensive sequence divergence, resulting in amino acid polymorphism within the A10-restricted epitope and a second A14-restricted epitope. Statistical analysis of the allelic sequences revealed evidence of positive selection for amino acid substitutions within the region encoding the CD8 T cell epitopes. Sequence differences in the A10-restricted epitope were shown to result in differential recognition by individual CD8 T cell clones, while clones also differed in their ability to recognize different alleles. Moreover, the representation of these clonal specificities within the responding CD8 T cell populations differed between animals. As well as providing an explanation for incomplete protection observed after heterologous parasite challenge of vaccinated cattle, these results have important implications for the choice of antigens for the development of novel subunit vaccines.
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spelling CGSpace1295962025-12-08T09:54:28Z Extensive Polymorphism and Evidence of Immune Selection in a Highly Dominant Antigen Recognized by Bovine CD8 T Cells Specific for Theileria annulata MacHugh, Niall D. Weir, William Burrells, Alison Lizundia, Regina Graham, Simon P. Taracha, Evans L. Shiels, Brian R. Langsley, Gordon Morrison, W. Ivan selection theileria polymorphism cells theileria annulata bovine Although parasite strain-restricted CD8 T cell responses have been described for several protozoa, the precise role of antigenic variability in immunity is poorly understood. The tick-borne protozoan parasiteTheileria annulatainfects leukocytes and causes an acute, often fatal lymphoproliferative disease in cattle. Building on previous evidence of strain-restricted CD8 T cell responses toT. annulata, this study set out to identify and characterize the variability of the target antigens. Three antigens were identified by screening expressed parasite cDNAs with specific CD8 T cell lines. In cattle expressing the A10 class I major histocompatibility complex haplotype, A10-restricted CD8 T cell responses were shown to be focused entirely on a single dominant epitope in one of these antigens (Ta9). Sequencing of the Ta9 gene from field isolates ofT. annulatademonstrated extensive sequence divergence, resulting in amino acid polymorphism within the A10-restricted epitope and a second A14-restricted epitope. Statistical analysis of the allelic sequences revealed evidence of positive selection for amino acid substitutions within the region encoding the CD8 T cell epitopes. Sequence differences in the A10-restricted epitope were shown to result in differential recognition by individual CD8 T cell clones, while clones also differed in their ability to recognize different alleles. Moreover, the representation of these clonal specificities within the responding CD8 T cell populations differed between animals. As well as providing an explanation for incomplete protection observed after heterologous parasite challenge of vaccinated cattle, these results have important implications for the choice of antigens for the development of novel subunit vaccines. 2011-05 2023-03-10T14:40:48Z 2023-03-10T14:40:48Z Journal Article https://hdl.handle.net/10568/129596 en Open Access American Society for Microbiology MacHugh, Niall D.; Weir, William; Burrells, Alison; Lizundia, Regina; Graham, Simon P.; Taracha, Evans L.; Shiels, Brian R.; Langsley, Gordon; Morrison, W. Ivan. 2011. Extensive Polymorphism and Evidence of Immune Selection in a Highly Dominant Antigen Recognized by Bovine CD8 T Cells Specific for Theileria annulata. Infection and Immunity 79: 2059-2069
spellingShingle selection
theileria
polymorphism
cells
theileria annulata
bovine
MacHugh, Niall D.
Weir, William
Burrells, Alison
Lizundia, Regina
Graham, Simon P.
Taracha, Evans L.
Shiels, Brian R.
Langsley, Gordon
Morrison, W. Ivan
Extensive Polymorphism and Evidence of Immune Selection in a Highly Dominant Antigen Recognized by Bovine CD8 T Cells Specific for Theileria annulata
title Extensive Polymorphism and Evidence of Immune Selection in a Highly Dominant Antigen Recognized by Bovine CD8 T Cells Specific for Theileria annulata
title_full Extensive Polymorphism and Evidence of Immune Selection in a Highly Dominant Antigen Recognized by Bovine CD8 T Cells Specific for Theileria annulata
title_fullStr Extensive Polymorphism and Evidence of Immune Selection in a Highly Dominant Antigen Recognized by Bovine CD8 T Cells Specific for Theileria annulata
title_full_unstemmed Extensive Polymorphism and Evidence of Immune Selection in a Highly Dominant Antigen Recognized by Bovine CD8 T Cells Specific for Theileria annulata
title_short Extensive Polymorphism and Evidence of Immune Selection in a Highly Dominant Antigen Recognized by Bovine CD8 T Cells Specific for Theileria annulata
title_sort extensive polymorphism and evidence of immune selection in a highly dominant antigen recognized by bovine cd8 t cells specific for theileria annulata
topic selection
theileria
polymorphism
cells
theileria annulata
bovine
url https://hdl.handle.net/10568/129596
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